Abstract
The hypothesis of an increase in free energy (exergy) by ecosystems during evolution is tested on direct measurements. As a measuring system of thermodynamic parameters (exergy, information, entropy), a series of measurements of reflected solar radiation in bands of Landsat multispectral imagery for 20 years is used. The thermodynamic parameters are compared for different types of ecosystems depending on the influx of solar radiation, weather conditions and the composition of communities. It is shown that maximization of free energy occurs only in a succession series (time scale of several hundred years), and on a short evolutionary time scale of several thousand years, various strategies of energy use are successfully implemented at the same time: forests always maximize exergy and, accordingly, transpiration, meadows-disequilibrium and biological productivity in summer, and swamps, due to a prompt response to changes in temperature and moisture, maintaining disequilibrium and productivity throughout the year. On the basis of the obtained regularities, we conclude that on an evolutionary time scale, the thermodynamic system changes in the direction of increasing biological productivity and saving moisture, which contradicts the hypothesis of maximizing free energy in the course of evolution.