Morphometric changes on dung beetle Dichotomius problematicus (Coleoptera: Scarabaeidae: Scarabaeinae) related to conversion of forest into grassland: A case of study in the Ecuadorian Amazonia

森林向草地转变对蜣螂 Dichotomius problematicus（鞘翅目：金龟科：蜣螂亚科）形态计量学的影响：以厄瓜多尔亚马逊地区为例