Studies in the biochemistry of micro-organisms: (a) On the production of hydroxyanthraquinones by species of Helminthosporium. (b) Isolation of tritisporin, a new metabolic product of Helminthosporium tritici-vulgaris Nisikado. (c) The molecular constitution of catenarin

微生物生物化学研究：(a) 蠕孢菌属真菌产生羟基蒽醌的研究；(b) 小麦蠕孢菌新代谢产物——小麦孢素的分离；(c) 链状素的分子结构研究。