Response Mechanism of Litter to Soil Water Conservation Functions Under the Density Gradient of Robinia pseudoacacia L. Forests in the Loess Plateau of the Western Shanxi Province

山西西部黄土高原刺槐林密度梯度下凋落物对土壤水分保持功能的响应机制

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Abstract

In the ecologically fragile western Shanxi Loess region, stand density regulation of artificial Robinia pseudoacacia L. forests plays a crucial role in sustaining the water regulation functions of the litter-soil system, yet multi-scale mechanistic analyses remain scarce. To address this gap, we established six stand density classes (ranging from 1200 to 3200 stems/ha) and quantified litter water-holding traits and soil physicochemical properties. We then applied principal component analysis (PCA) and structural equation modeling (SEM) to examine density-litter-soil relationships. Low-density stands (≤2000 stems/ha) exhibited significantly higher litter accumulation (6.08-6.37 t/ha) and greater litter water-holding capacity (maximum 20.58 t/ha) than the high-density stands (p < 0.05). Soil capillary water-holding capacity decreased with increasing density (4702.63-4863.28 t/ha overall), while non-capillary porosity (5.26-6.21%) and soil organic carbon (~12.5 g/kg) were higher in high-density stands (≥2800 stems/ha), reflecting a structural-carbon optimization trade-off. PCA revealed a primary hydrological function axis with low-density stands clustering in the positive quadrant, while high-density stands shifted toward nutrient-conservation traits. SEM confirmed that stand density affected soil capillary water-holding capacity indirectly through litter accumulation (significant indirect path; non-significant direct path), highlighting the central role of litter quantity. When density exceeded ~2400 stems/ha, litter decomposition rate decreased by ~56%, coinciding with capillary porosity falling below ~47%, a threshold linked to impaired balance between water storage and infiltration. These findings identify 1200-1600 stems/ha as the optimal density range; in this range, soil capillary water-holding capacity reached 4788-4863 t/ha, and available phosphorus remained ≥2.1 mg/kg, providing a density-centered, near-natural management paradigm for constructing "water-conservation vegetation" on the Loess Plateau.

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